lz^y4 adults have a "spectacle" eye phenotype. Ectopic cell death is seen in third larval instar eye discs.

Homozygotes have a total lack of ommatidial structure in the eye. The eyes may have mottled pigmentation with black pock marks visible on the surface of the eye, or pigment may be dispersed as a ring around the edge of the eye ("spectacle" phenotype).

Mutants display locomotor activity rhythms with circadian periods, though with reduced penetrance (small sample size).

The antennal funiculus is reduced in size in hemizygous males. All large and small basiconic sensilla on the antennal funiculus are missing, and the number of trichoid sensilla on the antennal funiculus is reduced. The number of coeloconic sensilla on the antennal funiculus is normal. The density of the coeloconic and trichoid sensilla on the antennal funiculus is increased compared to wild-type.

Eyes are greatly reduced in size and have a smooth surface in homozygous flies. Red eye pigment is sparse and confined primarily to the margin of the eye. Tarsal claws are vestigial. The spermathecae and pars ovariae are absent in homozygous females.

lz^y4/+ females have abnormal or absent female accessory glands and tend to lack spermathecae.

RK1. spectacle; spermathecae absent; parovaria absent Eye colour: redder than lz^s. Similar to lz^s but eye color redder.