hyd^C017 homozygous spermatogenic cells do not develop into spermatocytes.

hyd^hs1/hyd^C017 and hyd¹⁵/hyd^C017 mutants both exhibit abnormalities in spermatogenesis, with hyd^hs1/hyd^C017 exhibiting a more severe phenotype than hyd¹⁵/hyd^C017 mutants. One such abnormality is for nuclei scattered through the elongated cysts to exhibit a round shape instead of a 'needle' shape. In rare cases, the group of needle-shaped nuclei can be found in the correct place in the cyst; however, some of the nuclei are evidently incorrectly condensed. Approximately 90% of nuclei and micronuclei do not reach the correct cyst pole.

hyd¹⁵/hyd^C017 mutant spermatocytes exhibit defects in cytokinesis such as the formation of two spindles in one meiotic cell. The meiotic products may have unequal amounts of chromatin or mitochondrial material. Centrosome behavior is also affected. Meiotic chromosomes are not correctly attached to the spindle in 50% of hyd¹⁵/hyd^C017 spermatids.

hyd^C017 is a non-enhancer of visible phenotype of Scer\GAL4^GMR.PF, cindr^{RNAi.PC.PD.UAS}

hyd^C017 is a non-suppressor of visible phenotype of Scer\GAL4^GMR.PF, cindr^{RNAi.PC.PD.UAS}

hyd^C017 is a non-enhancer of eye phenotype of Scer\GAL4^GMR.PF, cindr^{RNAi.PC.PD.UAS}

hyd^C017 is a non-suppressor of eye phenotype of Scer\GAL4^GMR.PF, cindr^{RNAi.PC.PD.UAS}