Because of the In(1)wm4/wa constitution, C(1)M4 females without a Y chromosome display strong variegation, those with YS moderate variegation and those with YL or a complete Y almost no variegation. A powerful enhancer of autosomal recombination, but has low viability in combination with autosomal rearrangements.
C(1)M4, In(1)wm4 + AB - FM7., y wm4 - y- wa vOf sc8..