The wings of Doa⁴/Doa^γ3A mutant flies have blackened veins.

In Doa⁴/Doa^γ3A mutants where the Doa⁴ allele is from the mother the flies have badly oriented bristles on the head, missing or duplicated dorso-central bristles, forked scutellar bristles, fragile wings with incomplete posterior crossveins and darkened areas within the veins. When the Doa^γ3A allele comes from the mother there are no survivors.

In Doa^RemγA/Doa^γ3A mutants where the Doa^RemγA allele is from the mother the flies have rough eyes, missing dorso-central of scutellar bristles, slightly curly wings that are slightly translucent with small ectopic veins, generally posterior to wing vein L2, and disorganisation of the vaginal bristles. When the Doa^γ3A allele comes from the mother there are no survivors.