1 day memory retention is normal after spaced training in an olfactory conditioning assay.
The rhythmic change in bouton size that is seen in motor neuron MN5 under LD (12 hour light:12 hour dark) conditions and DD (constant darkness) conditions in wild-type flies is abolished in mutant flies.
Motor neuron MN5 shows increased neuronal branching in mutant flies compared to wild type.
Homozygotes are arrhythmic for locomotor activity under free-running conditions.