A short egg phenotype is seen as a result of disrupted cytoplasmic dumping from the nurse cells into the oocyte and result in eggs ranging from slightly shorter than wild-type to a quarter of the expected length. In some eggs the dorsal ventral polarity is affected, resulting in eggs which have fused appendages to various degrees or no appendage at all. When the cellularisation of mutant eggs is examined, it is sometimes not even across the entire surface and some eggs produce abnormal blastoderms. Abnormal gut invagination is also a common phenotype; invagination begins too far forward and towards the posterior and large 'holes' appear in their central yolk region. These embryos gradually become more abnormal as development proceeds and eventually generate without secreting cuticle. Other abnormalities are observed, where midgut invagination does not occur properly and also embryos are observed with abnormal head regions that fail to show any sign of head invagination.