Transheterozygotes are female sterile, exhibit no cuticular defects, meiosis is mostly normal with some nondisjunction and but do have brain defects. Heterozygotes with Df(3R)Hdγ1 are viable but female sterile.
Transheterozygotes with hsk1 are viable but female sterile. Transheterozygotes with cleo1 are lethal. Transheterozygotes with noose3 exhibit bristle duplication.
Phenotype assayed in trans with a multiple mutant chromosome carrying mutations of polo, mgr, asp, stg and gnu.