Embryos have completely unpigmented and weak cuticles. A portion of the homozygous embryos die before blastoderm formation with a defect in nuclear division (fewer nuclei that are misshapen, asynchronous and unevenly distributed), homozygous embryos that develop after have a segmentation defect (segment fusions and deletions).
complementing (Class IVc)
GTP cyclohydrolase activity is reduced in the prepupa, adult body and adult head of heterozygotes.
Class IV Pu allele.
Class IV allele.