UASp regulatory sequences drive expression of full-length cno isoform PE, mutated to carry amino acid replacements Y1553F, Y1565F, Y1652F, Y1679F, Y1727F, Y1800F, Y1875F, Y1927F, Y1932F, Y1948F, and Y1987F. The coding sequence is tagged at the N-terminal end with msVenus.