Adult expression is observed in ring neurons that arborize in the posterior disc and inner domain of the ellipsoid body.
In late third instar larva, expressed in the eye disc posterior to the morphogenetic furrow, in cells in the central brain and ventral ganglion, in specific regions of the wing and leg discs. In ovary, expression in border cells and follicle cells over the oocyte.
In ovaries, expressed in all follicle cells, including border cells.