Homozygous males are fertile, but their progeny show paternal chromosome loss (loss rate of paternal chromosome 4 is 21%). The timing of the chromosome loss is meiotic.
Crossover frequency in the X euchromatin (pn-f interval) is reduced 65-fold in soloZ2-3534 ordZ5736/soloZ2-0198 ord5 females compared to doubly heterozygous sibling controls. This reduction is intermediate in value between those of ord or solo single mutants.