Intergenic region between Dgri\Yp2 and Dgri\Yp1, extending to +10bp of Dgri\Yp2 and +40bp of Dgri\Yp1 is placed between lacZ and Ecol\CAT coding regions, such that the Dgri\Yp2 promoter is adjacent to lacZ and the Dgri\Yp1 promoter is adjacent to Ecol\CAT.