Mutant larvae that survive to the third instar stage empty their gut at the end of the wandering period, but do not contract or form a tanned puparium. Contraction of the peripodial epithelium, which allows the imaginal discs to evert in wild-type animals, does not occur in mutants, and the uneverted discs continue to elongate within vesicles inside the body. Muscles are present at the time of mouth hook detachment and retain their birefringence. Larval gastric caecae have shortened but are still present at the time of developmental arrest.