Embryos have completely unpigmented and weak cuticles. A portion of the homozygous embryos die before blastoderm formation with a defect in nuclear division (fewer nuclei that are misshapen, asynchronous and unevenly distributed), homozygous embryos that develop after have a segmentation defect (segment fusions and deletions).
Eye colour: mutant in trans with a homozygous viable Pu allele.
GTP cyclohydrolase activity is reduced in the adult head of heterozygotes. The viability and eye phenotype in trans with other Pu alleles has been determined.
Class IV Pu allele.
Class IV allele.