Embryos have completely unpigmented and weak cuticles. A portion of the homozygous embryos die before blastoderm formation with a defect in nuclear division (fewer nuclei that are misshapen, asynchronous and unevenly distributed), homozygous embryos that develop after have a segmentation defect (segment fusions and deletions).
complementing (Class IVc)
Class IV Pu allele.
Class IV allele.